Christina Skarpe & Johan T. du Toit 
Elephants and Savanna Woodland Ecosystems [EPUB ebook] 
A Study from Chobe National Park, Botswana

List of Contributors xi
Foreword xiii
Norman Owen-Smith
Preface xvii
Part I The Chobe Ecosystems 1
1. Introduction 3
Christina Skarpe and Stein R. Moe
2. The Chobe Environment 7
Christina Skarpe and Susan Ringrose
3. Elephant-Mediated Ecosystem Processes in Kalahari-Sand Woodlands 30
Johan T. du Toit, Stein R. Moe and Christina Skarpe
Part II The Substrate 41
4. Historical Changes of Vegetation in the Chobe Area 43
Christina Skarpe, Håkan Hytteborn, Stein R. Moe and Per Arild Aarrestad
5. Vegetation: Between Soils and Herbivores 61
Per Arild Aarrestad, Håkan Hytteborn, Gaseitsiwe Masunga and Christina Skarpe
Part III The Agent 89
6. Guns, Ivory and Disease: Past Influences on the Present Status of Botswana’s Elephants and their Habitats 91
Mark. E. Vandewalle and Kathy. A. Alexander
7. The Chobe Elephants: One Species, Two Niches 104
Sigbjørn Stokke and Johan T. du Toit
8. Surface Water and Elephant Ecology: Lessons from a Waterhole-Driven Ecosystem, Hwange National Park, Zimbabwe 118
Simon Chamaillé-Jammes, Marion Valeix, Hillary Madzikanda and Hervé Fritz
Part IV Controllers 133
9. Soil as Controller of and Responder to Elephant Activity 135
Christina Skarpe, Gaseitsiwe Masunga, Per Arild Aarrestad and Peter G.H. Frost
10. Impala as Controllers of Elephant-Driven Change within a Savanna Ecosystem 154
Stein R. Moe, Lucas Rutina, Håkan Hytteborn and Johan T. du Toit
11. Buffalo and Elephants: Competition and Facilitation in the Dry Season on the Chobe Floodplain 172
Duncan J. Halley, Cyril Taolo and Stein R. Moe
Part V Responders 187
12. Plant-Herbivore Interactions 189
Christina Skarpe, Roger Bergström, Shimane Makhabu, Tuulikki Rooke, Håkan Hytteborn and Kjell Danell
13. Elephants and the Grazing and Browsing Guilds 207
Christina Skarpe, Stein R. Moe, Märtha Wallgren and Sigbjørn Stokke
14. Cascading Effects on Smaller Mammals and Gallinaceous Birds of Elephant Impacts on Vegetation Structure 229
Sigbjørn Stokke, Sekgowa S. Motsumi, Thato B. Sejoe and Jon E. Swenson
15. The Chobe Riverfront Lion Population: A Large Predator as Responder to Elephant-Induced Habitat Heterogeneity 251
Harry P. Andreassen, Gosiame Neo-Mahupeleng, Øystein Flagstad and Per Wegge
Part VI Elephants in Social-Ecological Systems 269
16. Human Dimensions of Elephant Ecology 271
Eivin Røskaft, Thor Larsen, Rapelang Mojaphoko, A. H. M. Raihan Sarker and Craig Jackson
17. Elephants and Heterogeneity in Savanna Landscapes 289
Johan T. du Toit, Christina Skarpe and Stein R. Moe
Index 299

Christina Skarpe is a Professor in Applied Ecology at the
Faculty of Applied Ecology and Agricultural Sciences at Hedmark
University College, Norway. Her main research interest is large
herbivores and African savanna ecology. From Uppsala University,
Sweden, Botswana Ministry of Agriculture and later from Norwegian
Institute of Nature Research and Hedmark University College she
concentrated on herbivores- and worked on diverse projects in
Africa, Scandinavia, China and Central America.
Johan du Toit is Professor of Ecology and Conservation of
Large Mammals in the Wildland Resources Department at Utah State
University, USA. He has almost 30 years’ experience
conducting ecological research in African savanna ecosystems. At
the University of Zimbabwe he coordinated the Tropical Resource
Ecology Programme, after which he was the Austin Roberts Professor
of Mammalogy and Director of the Mammal Research Institute at the
University of Pretoria. His research focus is the ecology and
conservation of large mammals in terrestrial ecosystems.
Stein R. Moe is a Professor of Ecology at the Norwegian
University of Life Sciences, Norway. Although his scientific works
span over four continents, his main focus has been on African
savanna ecology. He is currently coordinating a Master’s
program in tropical ecology and natural resource management in the
Department of Ecology and Natural Resource Management. Following
several years as a departmental board member, he is now a member of
the faculty board at the University.